Tomarus Explained
Tomarus is a genus of scarab beetles in the subfamily Dynastinae, the rhinoceros beetles. They are native to the Americas, where they are distributed from the central United States to Argentina, and a few species occur in the Caribbean.[1]
The adult beetles are nocturnal and attracted to lights; the larvae generally remain in the soil, often feeding on plant roots.[1]
In order to identify species, the parameres of the male must be pulled out of the abdomen and examined. This process should be done carefully, because the parameres are quite fragile, "almost parchment-like".[1]
Species
The following species belong to the genus Tomarus:[1] [2]
- Tomarus adoceteus Ratcliffe & Cave, 2010
c g- Tomarus bidentulus (Fairmaire, 1892)
c- Tomarus bituberculatus (Palisot De Beauvois, 1805)
c g- Tomarus burmeisteri (Steinheil, 1872)
c g- Tomarus cicatricosus (Prell, 1937)
c g- Tomarus colombianus Lopez-Garcia & Gasca-Alvarez, 2014
g- Tomarus columbianus López-García & Gasca-Álvarez, 2014
c g- Tomarus compositus (Wickham, 1911)
c g
i c g b- Tomarus discrepans Escalona & Joly, 2006
c g- Tomarus ebenus (Degeer, 1774)
c g =
type species- Tomarus effetus (Wickham, 1914)
c g- Tomarus formosianus Grouvelle, 1913
g- Tomarus fossor (Latreille, 1833)
c g- Tomarus gyas Erichson, 1848
c g- Tomarus laevicollis (Bates, 1888)
i c g- Tomarus maimon Erichson, 1847
c g- Tomarus maternus (Prell, 1937)
c g- Tomarus nasutus (Burmeister, 1847)
i c g
i c g b
c- Tomarus pullus (Prell, 1937)
c g- Tomarus pumilus (Prell, 1937)
c g- Tomarus roigjunenti Neita & Ratcliffe, 2017
c g
c g- Tomarus rostratus Dupuis, 2014
c g- Tomarus rubripes (Boheman, 1858)
c g
i c g b- Tomarus selanderi (Cartwright, 1959)
i c g- Tomarus similis (Endrödi, 1968)
c g- Tomarus spinipenis Neita & Ratcliffe, 2017
c g
i c g b (sugarcane grub)
[3] - Tomarus villosus (Burmeister, 1847)
c gData sources: i = ITIS, c = Catalogue of Life, g = GBIF, b = Bugguide.net
- Note - now placed in the restored genus Ligyrus:
Notes and References
- Ratcliffe, B. C. and R. D. Cave. (2010). The Dynastinae (Coleoptera: Scarabaeidae) of the Cayman Islands (West Indies), with descriptions of Tomarus adoceteus, new species (Pentodontini) and Caymania nitidissima, new genus and species (Phileurini). Insecta Mundi 139, 1-15.
- Ratcliffe, B. C. (2003). The dynastine scarab beetles of Costa Rica and Panama (Coleoptera: Scarabaeidae: Dynastinae). Bulletin of the University of Nebraska State Museum 16, 1-506.
- Buss, E. A. Sugarcane Grub, Tomarus subtropicus Blatchley (Insecta: Coleoptera: Scarabaeidae). EENY-318 (IN593). Entomology and Nematology. Florida Cooperative Extension Service. University of Florida IFAS. Published 2003, revised 2006.